Nakia Cullain first/co-authored publications
Carpenter, M., Cullain, N., Daly, R., Smith, G., Dicken, M., Dames, V., Griffiths, C.L. and Marshall, A.D., 2025. Preliminary insights into manta ray (Mobula alfredi and M. birostris) population demographics and distribution in South Africa. Environmental Biology of Fishes, pp.1-23.
Carpenter, M., Cullain, N., Venables, S.K., Tibiriçá, Y., Griffiths, C. and Marshall, A.D., 2022. Evidence of Závora Bay as a critical site for reef manta rays, Mobula alfredi, in southern Mozambique. Journal of Fish Biology, 101(3), pp.628-639.
Cullain, N., McIver, R., Schmidt, A.L., Milewski, I. and Lotze, H.K., 2018. Potential impacts of finfish aquaculture on eelgrass (Zostera marina) beds and possible monitoring metrics for management: a case study in Atlantic Canada. PeerJ, 6, p.e5630.
Cullain, N., McIver, R., Schmidt, A.L. and Lotze, H.K., 2018. Spatial variation of macroinfaunal communities associated with Zostera marina beds across three biogeographic regions in Atlantic Canada. Estuaries and Coasts, 41(5), pp.1381-1396.
Cullain, N., Tibiriçá, Y., Venables, S.K., Rohner, C.A., Tittensor, D.P., Lotze, H.K., 2025. Declines in sightings and changing visitation patterns of reef manta rays at an important aggregation site in Mozambique. Environmental Biology of Fishes.
Marshall, A.D., Flam, A.L., Cullain, N., Carpenter, M., Conradie, J. and Venables, S.K., 2023. Southward range extension and transboundary movements of reef manta rays Mobula alfredi along the east African coastline. Journal of Fish Biology, 102(3), pp.628-634.
O’Connor, B. and Cullain, N., 2021. Distribution and community structure of at-risk and Data Deficient elasmobranchs in Zavora Bay, Mozambique. African Journal of Marine Science, 43(4), pp.521-532.
Van Driessche, C.D., Cullain, N., Tibiriçá, Y. and O’Connor, I., 2020. Fine-scale habitat use by humpback whales (Megaptera novaeangliae) in Zavora Bay, Mozambique. Western Indian Ocean Journal of Marine Science, 19(2), pp.131-147.
Marine Megafauna Foundation publications
Boggio-Pasqua, A., Flam, A.L. and Marshall, A.D., 2019. Spotting the “small eyes”: using photo-ID methodology to study a wild population of smalleye stingrays (Megatrygon microps) in southern Mozambique. PeerJ, 7, p.e7110.
Marshall, A.D. and Bennett, M.B., 2010. Reproductive ecology of the reef manta ray Manta alfredi in southern Mozambique. Journal of Fish biology, 77(1), pp.169-190.
Marshall, A.D., Compagno, L.J. and Bennett, M.B., 2009. Redescription of the genus Manta with resurrection of Manta alfredi (Krefft, 1868)(Chondrichthyes; Myliobatoidei; Mobulidae). Zootaxa, 2301(1), pp.1-28.
Pierce, S.J., White, W.T. and Marshall, A.D., 2008. New record of the smalleye stingray, Dasyatis microps (Myliobatiformes: Dasyatidae), from the western Indian Ocean. Zootaxa, 1734(6), pp.65-68.
Pierce, S.J., Méndez‐Jiménez, A., Collins, K., Rosero‐Caicedo, M. and Monadjem, A., 2010. Developing a Code of Conduct for whale shark interactions in Mozambique. Aquatic Conservation: Marine and Freshwater Ecosystems, 20(7), pp.782-788.
Rohner, C.A., Pierce, S.J., Marshall, A.D., Weeks, S.J., Bennett, M.B. and Richardson, A.J., 2013. Trends in sightings and environmental influences on a coastal aggregation of manta rays and whale sharks. Marine Ecology Progress Series, 482, pp.153-168.x
Rohner, C.A., Venables, S.K., Cochran, J.E., Prebble, C.E., Kuguru, B.L., Berumen, M.L. and Pierce, S.J., 2022. The need for long-term population monitoring of the world’s largest fish. Endangered Species Research, 47, pp.231-248.
Rohner, C.A., Venables, S.K., Knochel, A.M., Rambahiniarison, J.M., Marillac, V., Cardon, C., Scholten, N., Pierce, S.J. and Kiszka, J.J., 2025. Movements and habitat use of reef manta rays around the Mozambique Channel Island of Mayotte, Southwestern Indian Ocean. Environmental Biology of Fishes, 108(6), pp.937-955.
Venables, S., Winstanley, G., Bowles, L. and Marshall, A.D., 2016. A giant opportunity: the economic impact of manta rays on the Mozambican tourism industry—an incentive for increased management and protection. Tourism in marine environments, 12(1), pp.51-68.
Venables, S.K., Marshall, A.D., Armstrong, A.J., Tomkins, J.L. and Kennington, W.J., 2021. Genome-wide SNPs detect no evidence of genetic population structure for reef manta rays (Mobula alfredi) in southern Mozambique. Heredity, 126(2), pp.308-319.
Venables, S.K., Marshall, A.D., Germanov, E.S., Perryman, R.J., Tapilatu, R.F., Hendrawan, I.G., Flam, A.L., Van Keulen, M., Tomkins, J.L. and Kennington, W.J., 2019. It’s not all black and white: investigating colour polymorphism in manta rays across Indo-Pacific populations. Proceedings of the Royal Society B, 286(1912), p.20191879.
Venables, S.K., Rohner, C.A., Flam, A.L., Pierce, S.J. and Marshall, A.D., 2025. Persistent declines in sightings of manta and devil rays (Mobulidae) at a global hotspot in southern Mozambique. Environmental Biology of Fishes, 108(4), pp.749-765.
Venables, S.K., van Duinkerken, D.I., Rohner, C.A. and Marshall, A.D., 2020. Habitat use and movement patterns of reef manta rays Mobula alfredi in southern Mozambique. Marine Ecology Progress Series, 634, pp.99-114.